This page was partly financed by the Spanish Government through the research project EUI 2008-0388. Also by the Universidad Autónoma of Madrid (Spain) and the Regional Government (Comunidad de Madrid) through the research project CCG07-UAM/AMB- 1453.


The authors are also grateful to the curators of the cited herbaria for its kind assistance during ours visits and for loan of specimens.


Santiago Castroviejo


This website is dedicated to the memory of Dr. Santiago Castroviejo (1946-2009), friend and master.


Without his teachings, enthusiasm and unwavering support, this project, like many others, probably would not have seen the light.



Photo: Santiago Castroviejo in the Botanical Expedition to Serranía de Chiribiquete (Colombia) 1990


José María Cardiel, Elena Dorda, Pablo Muñoz & Manuel Pardo de Santayana
Dpto. de Biología (Unidad de Botánica), Facultad de Ciencias. UAM
Ciudad Universitaria de Cantoblanco. 28049 Madrid. España

Web development: Alfonso Marcos Vidal de la O & Marta Rubio Alfayate